Ursus arctos (carnivore)

Mass: 156 kg based on Kingsley et al. 1983

Diet: frugivore-carnivore based on Berns and Hensel 1972

Abundance: 0.01 to 36.13% (median 0.81%)

Latitudinal range: 30.3° to 68.8°

Habitats: (53), boreal forest/taiga (1), desert/xeric shrubland (1), montane grassland (4), temperate broadleaf/mixed forest (3), temperate coniferous forest (3), temperate savanna (2), tundra (1)

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Found in 68 samples

Canada: Grassy Bay

China: Suojia Township, Yanchiwan National Nature Reserve, Songhua River, Kanas National Nature Reserve

Croatia: Northern Velebit National Park

France: Les Canalettes (Layer 3), Les Canalettes (Layer 4), Coudoulous I (Faunal Unit II), Coudoulous I (Faunal Unit III), Coudoulous I (Faunal Units IV and V), Coudoulous I (Faunal Unit VI), Romain-la-Roche, Lazaret (C II supérieur UA 13-17), Payre (ensemble G), Payre (ensemble F), Payre (ensemble D), Tourville-la-Rivière (level D1)

Georgia: Dzudzuana Cave (lower area, Unit B)

Iran: Lar Protected Area, Wezmeh Cave, Golestan National Park

Israel: Marj Rabba

Italy: Western Trento Province (2012), Caverna degli Orsi (sondage A, Unit 11.2), Riparo Biarzo (SU 5), Riparo Biarzo (SU 4 + 3B), Riparo Biarzo (SU 3A), Mondeval de Sora (SU 8), Mondeval de Sora (SU 31), La Sassa Cave, Grotta Cucgiliana

Lebanon: Moghr el-Ahwal Cave 3 (Phases F & E)

North Korea: Chongphadae Cave

Poland: Borsuka Cave (Layer VI), Borsuka Cave (Layer III)

Portugal: Gruta de Figueira Brava (Area C)

Romania: Southern Transylvania, Noviodunum (TPF sector, Middle Ages)

Russia: Geographical Society Cave, Chagyrskaya Cave (Layer 5), Chagyrskaya Cave (Layer 6), Cheremukhovo-1 (Bed 4), Imanay Cave (Pleistocene Layer 1), Imanay Cave (Pleistocene Layer 2), Imanay Cave (Pleistocene Layer 3), Imanay Cave (Pleistocene Layer 4), Khayrgas Cave - Layer 6 (1998–1999 pit), Khayrgas Cave - Layer 7 (1998–1999 pit), Byzovaya

Spain: Cueva del Angel, Bolomor Cave (Level IV)

Tunisia: Meninx

Turkey: Yaylacik Research Forest, Marmaris, Barhal Stream, Direkli Cave

United Kingdom: Lynford Quarry, Balderton Sand and Gravel

United States: Dalton Highway, Welsh Cave, Wewukiyepuh, Polecat Creek (alluvial gravel), Schulze Cave (layer B), Schulze Cave (layer C1), Molybdenum Ridge (triggered detections), Molybdenum Ridge (time-lapse detections), Black Rapids Training Area (triggered detections)

Size measurements:
♀ body mass105000 kgN = 45Kingsley et al. 1983
♀ body mass198900 kgN = 19Kingsley et al. 1983
♀ body mass91000 kgN = 24Kingsley et al. 1983
♀ body mass162500 kgN = 18Kingsley et al. 1983
♂ body mass190600 kgN = 41Kingsley et al. 1983
♂ body mass258300 kgN = 10Kingsley et al. 1983
♂ body mass190400 kgN = 24Kingsley et al. 1983
♂ body mass222400 kgN = 19Kingsley et al. 1983

See also Ursus, Ursus americanus, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus, Ursus thibetanus, Ursus vitabilis